Scientists are increasingly focused on balancing fisheries management with long-term sustainability, a challenge addressed by new research modelling optimal harvesting strategies within marine protected areas. Dinesh Kumar, leading this work, investigates a two-patch model incorporating both harvestable and preserved zones to determine how best to maintain fish stocks and maximise revenue. This study is significant because it moves beyond simple constant-effort policies, demonstrating that a composite Bang-Singular-Bang control strategy , derived using Pontryagin’s Maximum Principle , can substantially improve both ecological and economic outcomes. Simulations reveal that even modest levels of preservation (20-30%) can support more intensive harvesting while preventing population collapse, offering valuable insights for fisheries management worldwide.
Dinesh Kumar, leading this work, investigates a two-patch model incorporating both harvestable and preserved zones to determine how best to maintain fish stocks and maximise revenue. Simulations reveal that even modest levels of preservation (20-30%) can support more intensive harvesting while preventing population collapse, offering valuable insights for fisheries management worldwide., Numerical simulations reveal this dynamic strategy?
Optimal Harvesting with Limited Marine Protection requires careful
Keywords: Marine protected area, Seasonal harvesting, Beverton, Holt recruitment, Bio-economic optimal control, Bang-bang control, Singular control, Pontryagin maximum principle. Marine fisheries represent a vital renewable resource, supplying essential animal protein to billions of people and supporting livelihoods for hundreds of millions worldwide. Yet, the history of fisheries management is punctuated by severe stock collapses, exemplified by the Atlantic cod fishery off Newfoundland in the early 1990s and the recent climate-driven declines in Pacific cod in the Gulf of Alaska. Such collapses entail profound ecological, economic, and social impacts, with recovery often proving slow and uncertain even after substantial reductions in fishing pressure.
At its core, the challenge for fisheries management is bioeconomic: balancing maximization of yields and economic benefits with the preservation of population viability and ecosystem integrity. Traditional fishery management, centred on maximum sustainable yield (MSY) principles, has been widely critiqued as inadequate for capturing the complex spatial, temporal, and multispecies dynamics of fish populations. Contemporary approaches increasingly incorporate spatial tools, such as marine protected areas (MPAs), particularly no-take reserves where extractive activities are prohibited. Empirical evidence and theoretical models suggest that well-designed MPAs can enhance both conservation and fishery outcomes through multiple mechanisms: providing refuges for spawning populations, protecting critical habitats, facilitating spillover of adults and larvae into adjacent fishing grounds, and reducing fishing mortality on juvenile cohorts.
The mathematical theory of renewable resource economics traces to the seminal work of, who identified the “tragedy of the commons” in open-access fisheries, and, who formalized the concept of maximum sustainable yield. pioneered the application of optimal control theory to fishery management, demonstrating that purely economic optimization can paradoxically lead to population extinction when economic discount rates exceed biological growth rates, the so-called “Clark’s rule”. His monograph remains the definitive reference for bioeconomic modelling, establishing the theoretical foundation for much subsequent work. Early models assumed homogeneous, non-spatial populations. Extensions incorporated age or stage structure, stochastic environmental variability, and Allee effects.
Spatial heterogeneity, however, gained attention later. pioneered spatially explicit bioeconomic models, analysing fishing behaviour across heterogeneous patches with dispersal. They showed that spatial variation in fishing costs or stock productivity can lead to de facto reserves emerging from economic optimization, even without regulatory intervention. derived the conditions under which marine reserves enhance fishery yields, identifying critical thresholds in dispersal rates and reserve sizes that determine whether reserves benefit or harm adjacent fisheries. Empirical support for marine protected areas (MPAs) grew via meta-analyses showing consistent increases in biomass, density, size, and diversity within reserves. Spillover to fisheries is more variable and context-dependent, underscoring the value of coupled reserve-fishery models.
Recent work has extended spatial models in several directions. analysed dynamic spatial closures, showing that adaptive management of reserve boundaries can substantially improve economic outcomes. incorporated spatial degradation and recovery dynamics, demonstrating complex trade-offs between conservation and economic objectives. reviewed ecological connectivity in MPA network design, emphasizing the importance of larval dispersal and adult movement patterns. Many commercially important fish species exhibit pronounced seasonal patterns, with distinct periods for spawning, juvenile development, and adult feeding/migration. Classical continuous-time models implicitly assume constant vital rates and overlook these temporal structures. Discrete-time models with seasonal components better capture species like Pacific salmon, Atlantic herring, and many tropical reef fish. analysed seasonal harvesting models with time delays, showing that harvesting during non-reproductive periods can sustain higher yields than year-round exploitation. demonstrated that migratory dynamics fundamentally alter optimal harvesting strategies, with implications for transboundary fisheries management.
Our model builds on this tradition by explicitly separating reproduction (occurring in non-fishing season and potentially in reserves) from harvesting (occurring in designated fishing zones during limited seasons). The Beverton-Holt recruitment function, which we employ, has been extensively validated for stocks exhibiting strong density-dependent recruitment at the larval or juvenile stage. This functional form arises naturally when early life stages experience contest competition or predation that saturates at high densities, contrasting with the Ricker model appropriate for scramble competition. applied PMP to fishery harvesting, deriving the classic result that optimal policies often exhibit “bang-bang” control, switching between maximum and zero effort, or “singular” control, where effort follows a state-dependent feedback rule.
Our model possesses Markov structure through its seasonal discrete-time formulation: the population state at the beginning of each harvesting season fully determines future dynamics given control choices. This enables decomposition of the infinite-horizon problem into a sequence of single-season optimizations, each solved via PMP, with the Bellman equation providing the linking condition across seasons. In this work our contributions are: a mathematical model: a two-patch system coupling within-season continuous dynamics (mortality, dispersal, harvesting) with discrete-time seasonal reproduction via the Beverton-Holt function. One patch is a no-take reserve; the other allows controlled harvesting. stability analysis: explicit derivation of the condition Fr > 1 for population persistence, where F encapsulates within-season survival (including harvesting effects) and r is intrinsic growth.
Complete characterization of transcritical bifurcation surfaces in (T, R, E) parameter space, showing how preservation zones expand stability regions. Proof that controls follow bang-singular-bang structure. Derivation of explicit state-feedback formula for singular arcs and verification of GLC optimality conditions. We assume a spatially separated population with a harvesting area and a no-take reserve. The dynamics of the population are described by the following system of equations: x1x2 = −c −qE −mR m(1 −R) mR −c −m(1 −R) x1x2, where x1 and x2 represent the population density in the harvesting and reserve areas, respectively. c is the natural mortality rate, q is the catchability coefficient, E is the fishing effort, m is the dispersal rate between the two areas, and R is the proportion of the population in the reserve.
Rewriting the system in vector form, X = AX, where X = x1x2 and A = −c −qE −mR m(1 −R) mR −c −m(1 −R). The solution of the system is given by X(t) = eAtX0, where X0 is the initial condition. Thus, X(k) = eAT X(k −T), where T is the duration of the fishing season. Reproduction is modelled using the Beverton-Holt function: x(k) = rx(k −1) 1 + βx(k −1). This function represents the growth rate r and the density effect coefficient β.
Combining the system dynamics and the reproduction function, we obtain the discrete. The eigenvalues λi are given by λi = −T 2 2c + m + qE + (−1)i+1p (m −qE)2 + 4mqER, (i = 1, 2). The equilibrium points of the discrete system are 0 and α−1 β, α > 1. Long-term sustainable harvesting is possible only when α > 1, i. e., Fr > 1.
Protected areas boost sustainable fishery viability by safeguarding
Scientists have developed a bioeconomic model for optimal fishery harvesting within spatially heterogeneous environments, incorporating both harvestable and preservation zones. The population dynamics are governed by a hybrid system, coupling continuous time within-season dynamics, mortality, harvesting, and dispersal, with a discrete-time Beverton-Holt reproduction map. The simulations showed a stable “sawtooth” trajectory, indicating a sustainable balance between harvesting and population growth. This work highlights the potential for integrating preservation strategies with economic optimisation in fisheries management.
The research demonstrates that a carefully designed balance between protected and harvested areas can enhance both ecological sustainability and economic returns. Specifically, the model predicts that implementing marine protected areas can expand the range of parameters under which a fish population can persist, offering a buffer against environmental fluctuations and overfishing. The derived control strategy provides a framework for adaptive management, allowing fisheries managers to respond to changing conditions and optimise harvesting practices for long-term sustainability.
Marine Reserves Boost Fisheries Sustainability Parameters significantly
Scientists have developed a bioeconomic model to determine optimal fishery harvesting strategies in spatially complex environments, incorporating both harvestable areas and marine reserves. The model integrates continuous dynamics representing within-season population changes with a discrete-time reproduction map, allowing for analysis of long-term population persistence under varying harvesting pressures. Researchers derived a condition for persistence, demonstrating that a sufficient level of within-season survival, influenced by harvesting, is crucial alongside the intrinsic growth rate of the population. The costate variables within the model represent shadow prices, indicating the marginal value of fish in each zone and justifying aggressive late-season harvesting, provided minimum viable populations are maintained.
This research provides a theoretical basis for ecosystem-based fisheries management, demonstrating that ecological and economic objectives can be aligned through strategic spatial refuges and dynamic control measures. The authors acknowledge limitations including the absence of stochasticity, age structure, and multi-species interactions within the current model. Future research will focus on incorporating these complexities to further refine the policy relevance of the findings and enhance the model’s predictive capabilities.
👉 More information
🗞 Sustainable and Optimal Harvesting in a Seasonally Harvested Fishery with a Marine Protected Area: A Two-Patch Model with Bang-Bang and Singular Control
🧠 ArXiv: https://arxiv.org/abs/2601.20563
